This thesis investigates the subsistence behaviour of Early Upper Palaeolithic (EUP) Homo sapiens based on the remains of molluscs and terrestrial fauna recovered from the Ksâr ‘Akil rockshelter ... Show moreThis thesis investigates the subsistence behaviour of Early Upper Palaeolithic (EUP) Homo sapiens based on the remains of molluscs and terrestrial fauna recovered from the Ksâr ‘Akil rockshelter (Lebanon). The results tie into the debate around the dispersal of modern humans into Europe. In this thesis, a multi-proxy chronological approach was applied providing ages for the modern human occupation and behavioural adaptations just outside Europe. At the onset of the Upper Palaeolithic groups at Ksâr ‘Akil relied mainly on terrestrial food resources, which shifted later, during the early Ahmarian, gradually to more extensive exploitation of smaller bodied taxa including the introduction of marine and terrestrial snails into the diet. Oxygen isotope analysis of marine mollusc taxa showed that shellfish exploitation was practiced in all seasons and thus played a central role in EUP foraging strategies. Equally this implies that the site was occupied during different times of the year. With regard of the implications for EUP Homo sapiens, shellfish as a dietary supplement throughout the year enhances the diet nutritionally and likely contributes to healthier populations. This in turn might have facilitated population growth and increased population density in an area that fuelled the modern human dispersals into Europe. Show less
Palaeoanthropologists have proposed that Neanderthals, the Middle Palaeolithic hominin occupant of Eurasia, differed from modern human relatives by having specialised diets focused on big game. A... Show morePalaeoanthropologists have proposed that Neanderthals, the Middle Palaeolithic hominin occupant of Eurasia, differed from modern human relatives by having specialised diets focused on big game. A narrow dietary niche at the top of the terrestrial food chain is inherently prone to instability, potentially contributing to extinction of the Neanderthals. However, limitations in detecting plant consumption imply that scientists are unaware of much of Neanderthal diet. My dissertation revises the role of plants in Neanderthal subsistence using dental calculus, a material that is recognised to contain food traces, as a source of dietary data. To do this I assessed how accurately calculus records diet, by testing it with a variety of techniques on a population of chimpanzees with decades of documented dietary history. Then, my dissertation examined if it is possible to explore the resilience of the Neanderthal dietary niche by assessing for changes in plant use over time. Comparing diets from different habitats, data suggests a broad range of diets on the Mediterranean rim and in the cooler areas of the Neanderthal range. Surprisingly, the study found no evidence of changes in plant dietary breadth despite variation in environments. This stability implies a deeply resilient ecological niche across their range. Show less